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All males from 20 years upward who were able to go to war; this excluded the infirm, aged, and maimed.

To do the service of the Tabernacle; to minister unto Aaron and his sons.

Every male from a month old and upward.

Merarites, Gershonites, Kohathites, Eleazar, and Ithamar.

The male first-born; the first-born were chosen as Hashem's special ones in Egypt when the the first born of all the Egyptians were killed. When the first-born sinned by participating in the worship of the golden calf, they were replaced by the Levites who received their special priestly privileges.

The first 22,000 were redeemed, each one according to one of the 22,000 Levites. The extra 273 first born that had no corresponding Levite, had to pay 5 shekels each to be redeemed; The Levites did not have to be redeemed.

Moshe gave the redemption money to Aaron and his sons as commanded by Hashem.